Lasallia pertusa (Rassad.) Llano
  Monograph of the Umbilicariaceae 48 (1950).
  TYPE: Russia, Siberia, LakeBaikal, western lakeside, between 53<-54< Lat. s., Valey Onguren, on boulders in leaf-bearing forest, July 10, 1928, coll. K.A.Rassadina No. 289 (LE-lectotype!).
  《 Umbilicaria pertusa Rassad. Comp. Rend. Acad. Sci. URSS, 14 (A): 348 (1929), fig. 1-2.
  《Lasallia pensylvanica var. pertusa (Rassad.) Llano, Hvalrad Skr. 48: 122 (1965).
  TYPE OBSERVATION:Four syntype specimens (Nos. 25, 119, 289) cited by Rassadina in her original description are preserved in LE. All of these four specimens were available for this study, and corresponding completely in their characteristics to the original description. One of these four specimens(No. 289) is selected here as a lectotype for this species.
  Thallus monophyllous, up to 7-9cm diam, perforated, perforation on the top of the pustules; perforation margines sorediose with few isidia; upper surface areolate-reticulate, brownish; lower surface light coloured, black around the umbo, margins whitish to greyish, lacunose.
  Apothecia unknown.
  =Lasallia rossica var. pertusoides Dombr. Nov. syst. plant. non vascul. 15: 186(1978).
  Type: Russia, districtus nationalis czukotensis, systema fl. Belaja, affluxionis sinistrae fl. Anadyrj, in depositionibus jurassicis ad fl. Inmuam, inter Ustj-Chvuk et ostium fl. Muchomonaja, Aug. 23, 1929, coll. V.B.Soczava(LE!).
  =Lasallia pertusa f. squamulifera Wei & Jiang, Acta Phytotax. Sin. 20(4): 499(1982).
  Type: China,Xizang, Lhasa, on rocks, alt. 3750m, April 3, 1966, coll. Wei JC & Chen JB, No. 15(HMAS-L, holotype!); Gyrong, alt. 2880m, Jun. 14, 1975, coll. Zong YC, No. 112(HMAS-L, paratype!).
  DESCRIPTION: Thallus monophyllous, orbicular, c.4cm diam., with abundant fenestrate pustules, soredia, and few isidia; upper surface finely areolate, grey-brown red- brown to dark brown, dull to slightly shining; lower surface greyish brown, covered with a thin layer of white pruina, dark brown to black-brown around the umbo.
  Thallus 371-653μm thick; necral zone 7-18μm thick; upper cortex paraplectenchymatous, 35-46μm thick, including a brown outer layer of 7-14μm thick and a colourless inner layer 28-32μm thick; algal layer continuous, 48-130μm thick; medulla mucilaginous, 195-413μm thick; lower cortex 65-107μm thick, including a colourless inner layer 18-37μm thick, and a grey to brownish outer layer 37-70μm thick.
  Apothecia not seen.
  Pycnidia, when immature, 93-185〜139-255μm. No pycnoconidia seen.
  CHEMISTRY: lecanoric and gyrophoric acids, with an orange-yellow pigmentin Rf class 3 if B solvant.
  SPECIMENS EXAMINED: 
  China: Xinjiang, by the river near Buljin tree farm, alt. 1300m, Sept. 13, 1982, coll. Zhao CF No. 2458(HMAS-L!). Xizang, Nyalam, on rocks, alt. 3700-3720m, April 29, & May 11,15, 1966, coll. Wei JC & Chen JB, Nos. 92, 1429, 1648(HMAS-L!).
  Himalaya: Pangi, Coll. D.Scoliczha(US!).
  Mongolia:Ara-Khangai, on rocks, July 17, 1970, coll. Biazrov. No. 6685(MOSCOW!) on rocks, July 13, 1977, coll. Biazrov, No. 3300(MOSCOW!); on rocks, Aug. 21,22,1979, coll. Biazrov, Nos. 6586, and on bark of Larix sibirica: Nos. 4384, 6587,6589,6599 (MOSCOW!);July 1, 1980, on bark of Salix sp., coll. Biazrov, No. 4325(MOWCOW!); Ju- ly 21, 1980, coll. Biazrov,No. 4383(MOWCOW!)Bulgan, onBetula sp., Jun. 29, 1972, coll. Biazrov, No. 5088(MOSCOW!); on rocks, Jun. 29, 1972, coll. Biazrov, Nos. 3280, 3293, 5039(MOSCOW!). Chentyi, valley of Churchingol river, in mt. Chutag-Chayran ula, on rocks in Larix sibirica forest, July 15, 1974, coll. N. Golubkova & Zogt, No. 2056 (LE!). Chobgol, Aug. 3, 1972, coll. Golubkova & Zogt, No. 681(LE!); Western Mongolia Aug. 2, 1972, coll. Zogt.No. 606(LE!)Zabhan, on roks, Jun. 29, coll. Biazrov, No. 6585 (MOSCOW!); on rocks; July 4,11, 1976, coll. Biazrov, Nos. 6581, 6583(MOSCOW!).
  Russia: Siberia, Lake Baikal, western lakeside, between 53≧ and 55<Lat. S., Island Olkon, on rocks, Jun. 18, 1928, coll. K.A. Rassadina, No. 119(LE! syntype); In lapidosis Daburiae 1832, herb. H.Kuznetzov. Herb. Nos. 25, 26 bearing Umbilicaria pustulata(LE! syntype). Krasnojarsk, lat. bor. 56<Jun. 11, 1876, M. Brenner. No. 302f, as U. pensylvanica (S!)Minusinksk, on rocks, (July?)Aug. 1901, coll. Martianoff(H, TUR!), & July 24, 1926, coll. W. Reverd atta No. 1(H!)Regio Irkutensis, ad oram Occidentalem lacus Bajkal, ad saxa in sylva laricina et in insula Olchon ad rupes. 1928, coll. K.A.Rassadina No. 51 as U. pensylvanica f. pertusa Rassad. (MIN!).Gouv Tenisejeu, Slus Tenosej Auf Gestein Jun. 30, 1926, coll. W.Renerdotti, No. 33(FH!). SW Siberia, Altai Mts., Gorno-Altaisk Autonom. Oblast. SE of Ongudai on little Ilzumen River near Khabarovka, below Chikemansky Pass, 900-1000m.s.m., arid mtn., Aug 20,1978, coll. T.S.Elias, W.Weber, C.S.Tomb & I.M. Krasnoborov, as U. pustulata(COLO!).
  Nepal: Central Himalaya, Langtang, alt. 3600m, Sept. 8, 1896, coll. J.Poelt, L-84318, C.H.Colo-411988(COLO!).
  LITERAURE RECORDS FOR ASIA: 
  China(Wei JC & Chen JB, 1974, p. 177; Wei JC, 1981,p.90; Wei JC & Jiang YM, 1981, p. 1146 & 1986,p.102).
  Mongolia(R.Schubert & O.Klement, 1971, p. 226; Golubkova, 1981, p.134; Huneck et al., 1992, p.296).
  Russia(Rassadina, 1929, p.348; Krog, 1973, p.553; Golubkova et al., 1978, p. 131 for Siberia; Sedelnikova, 1985, p.89 for Siberia; Makrei, 1990, p. 55 for Siberia.).
  Himalaya(Llano, 1950, p.48; Awasthi, 1965, p.125, as U. pertusa; Frey & Poelt, 1977, .389).
  COMMENTS: This species is very similar to L. papulosa, but differs in the fenestrate pustules and in having soredia on the tips of the pustules or along the perforated rim. 6.Lasallia pustulata(L.)Merat(Figs.24-26; map 1) Nouv. Fl. Paris, ed. 2,1:202(1821).